Monday, 28 August 2017

Romania - Southern Transylvania & Southern Carpathians

Southern Transylvania

The region between Sibiu & Sighisoara holds a series of north-south directed valleys in a hilly landscape with extensive grasslands in the lower parts of the valleys and forests on the hilltops. We visited 2 of those valleys, one near Biertan and one just SW of Sighisoara.

Iphiclides podalirius

Hamearis lucina, having a second generation in central Europe, unlike in western Europe where this is mainly a spring species.

Lycaena tityrus

Phengaris arion

Phengaris teleius on the larval foodplant Sanguisorba officinalis

Plebejus argus

Neptis sappho, much more common than Neptis rivularis (see previous post) but this species has 2 generations while N. rivularis only has one. During our trip we were probably largely too late for N. rivularis, except for the ones flying at higher elevations.

Argynnis adippe, a nice variation of the cleodoxa colour morph with the submarginal band accentuated.

Star of this part is Argynnis laodice, photography wasn't easy however as the favorite way to spend  time of these Argynnis species is to attack any other Argynnis (here A. laodice is attacked by A. adippe). So when you saw one landing on these big Telekia flowers mostly you had some 10 seconds before it was chased away... and A. laodice was outnumbered by A. paphia & A. adippe.

Argynnis laodice

Southern Carpathians
We visited two of the several southern Carpathian mountain chains. In the Făgăraş Mountains I searched for butterflies on the southern slopes between 1550m & 1950m. In Bucegi Mountains between 1400m & 2000m. Unlike the Alps the Carpathians are rather poor in butterlfy diversity and on a day high up in the mountains it was not rare to come back with a species list of only 10-15 butterfly species.

Boloria pales carpathomeridionalis, the most fun about this subspecies is that Boloria napaea is absent in the Carpathians so there is no determination issue. Several seen in Bucegi mountains.

 Erebia epiphron transsylvanica, by far the most common butterfly on the grasslands high in the Carpathians, both in the Făgăraş Mountains as in the Bucegi Mountains. 

Erebia euryale syrmia. Commonly present on grasslands near the pine forest zone. The look-a-like Erebia ligea was present at a lot of locations as well but mostly a bit lower, on open places in the forest zone.

Erebia manto trajanus, only one very fresh male seen at the Făgăraş Mountains, suggesting that due to the harsh Eastern European winter the flight period of this species had only just begun at the end of july (30/07/2017). Unfortunately this male wasn't very happy with me trying to get it on camera, flying uphill very fast after this one record shot...

Erebia pronoe regalis, several seen on the higher slopes of Bucegi Mountains (1700-2000m).

Erebia sudetica radnaensis at the Făgăraş Mountains. This subspecies has only very faint eyespots on the upperside, they just fall in the folds of the veins so almost seem absent. The same remark as with Erebia manto, only one fresh male seen so probably only the very beginning of the flight period. More information on this species in Romania can be found here.

From the mountains to the plains and marshes in the next episode!

Romania 2017 - Introduction

Last month I travelled through Romania, a country I already wanted to visit for a long time. We entered the country the 25th of july & left it again on 12th of august.
Except for the first & the last day we had splendid weather, at some days even too hot to watch butterflies on a comfortable way. The bad weather on first & last day were a bit pitty as that were the days we passed the region around Cluj-Napoca and there were some locations in that neighbourhood I wanted to visit. Luckily I didn't really miss a big target species because of that. Time was not only spent on butterflies on this trip, I also looked a bit at birds & dragonflies.

We went by car from Belgium entering Romania from the west near Oradea. The locations we visited can be resumed in the folowing 8 points:

1) Turda gorge & Piatra Secuiului (Eastern slopes of Apuseni Mountains)
Calcareous hills & rocky outcrops between 700m-1200m

2) Southern Transylvania
Lower hills with lots of extensive grasslands & decideous forest

3) Fagarasan Mountains (Southern Carpathians)
High mountains up to 2500m

4) Bucegi Mountains (Southern Carpathians)
High mountains up to 2500m

5) Plains S of Danube
Low coastal plains, mainly visited to see dragonflies & birds but with some interesting butterfly sightings

6) Danube delta
Marsh land with canals, lakes, willow forests

7) Babadag forest & Macin mountains
Low mountains with dry oak forests

8) Foothills of the eastern Carpathians
Extensive grasslands 700-1000m




So let's kick off this small report with some pics

1) Turda gorge & Piatra Secuiului
Turda gorge is a pretty spectacular 300m deep gorge but from a butterfly point of view especially the plains up the gorge are interesting.

Habitat in Turda


Hesperia comma is common in large parts of Romania, very different from the Western part of the distribution where this species is on many locations threatened.

Dark Hipparchia's are easy in Romania, in Transylvania & the foothills of the Carpathians only Hipparchia fagi is present, in Dobrodgea probably only Hipparchia syriaca is present. This is Hipparchia fagi.

Iphiclides podalirius, although common in large parts of southern Europe, always a pleasure to see

As this one seems to have a discocellular vein, closing down the middle cell, this should be Melitaea trivia. Opinions welcome.

Polyommatus dorylas, male, rather common in this part of Romania

Pseudophilotes vicrama

We visited Piatra Secuiului to see the local Erebia melas. I saw a few of them flying and a single one shortly visiting some flowers but the habitat with very steep calcareous rocks didn't allow taking pictures, although I tried... In the neighbourhood however I did make a good set of pictures, definetely a blues paradise!

Habitat

Neptis rivularis, only a few seen during the trip and mostly higher up (this one at approx 1000m)

Polyommatus daphnis, male

Polyommatus daphnis, female

Polyommatus daphnis, female, one of the most marvelous species of blues

Polyommatus dorylas, male

Polyommatus thersites, male. Underside & upperside of the same individual, notice the hairy basal half of upper front wing. The hairy androconial scales are the single best feature to differentiate this species from Polyommatus icarus.

Cupido decoloratus, male

Cupido osiris, male

More on Transylvania in the next episode!



Sunday, 2 July 2017

SW-Germany (3): Nymphalids

Melitaea didyma
In the Moselle valley this species is present but I guess I just missed out the first generation (250m), in the Swäbische Alb however the species was fresh and common (800m).



Melitaea britomartis
My main target for this trip. During summer trips to the east I always missed out on this species being slightly too late. So I wanted to see them a bit more nearby and the closest to find them near Belgium are some grasslands in the submontane zone of the Swäbische Alb. As most of you know this species is in a group of species that is really difficult to recognise in the field (subgenus Mellicta, sometimes called the Melitaea athalia-group). Lot of the features mentioned in the fieldguides for these species are very variable individually making it very difficult to point an individual to a species (see for example this article)
Lafranchis gives a method in his guide to check genitalia of butterflies in the field. He gives this for several of groups of butterflies. 
Frankly I think for the Leptidea species his images are the worst you could ever use to see the difference between the cryptic Leptidea species and difference in genitalia between Leptidea sinapis and Leptidea reali/juvernica can't be seen in the field. Anyone using the Lafranchis guide should make a big black cross over the drawings of the Leptidea genitalia!
For the orange Hipparchia species as well I don't think this method is very reliable as often you have to make an interpretation of the scale of different parts of the valves which is very difficult in the field. In the black Hipparchia species you just have to count the bristles on the Julien organ what seems pretty easy in the field to me. 
In the Melitaea species the shape of the valves are clearly different between species so there the method leads to more reliable determinations too imo. You need some experience in handling the butterflies and you need some experience to evaluate some individual variation in the shape of the valves. The image of the M. britomartis valve in Lafranchis is not that good in my opinion, the image in Slamka however was spot on.
At the same location where I had these M. britomartis I had M. aurelia and M. athalia males! Mostly M. athalia was easily recognised in the field as they are a bit bigger and clearly have a more direct flight than the smaller M. britomartis and M. aurelia. Males of both these species have a very specific flight with fast wing beats making them stand out between the M. athalia males.

male underside

same individual, male upperside

Male genitalia checked in the field with the insert showing the valve of a genital preparation (edited picture from lepiforum). With a 20x loupe details of the valve shape can easily be seen.

Female, in front view this one has quite some black hairs on the palps, strongly indicating it not to be M. aurelia (variable in M. britomartis but probably almost always orange or at least pale in M. aurelia). The orangeish color in the marginal line is typical (but variable) for M. britomartis.


Argynnis niobe
This species used to be present in the Belgian coastal dunes as well as in the calcareous grasslands of the south but now has completely disappeared. It is still present however just south of Aachen, less than 10km of the Belgian border. Several individuals were present when I passed that location but photography was impossible, all seemed to be males actively searching for females and gathering around some old pine trees using them to do some kind of hill topping. I saw the species in the south of the Black Forest as well where I could make some pictures.



Boloria titania
I was pleasantly surprised to find this species in the Black Forest, the main field guides don't give this as a part of the distribution. Of course when one does a bit more research this is well known and the species is present just north up to Triberg (southern 1/3 of the Black Forest).




Coenonympha tullia
I had seen this species before but then was unable to make good pictures, so I was hoping to make some pictures on this trip. I found it in the same bog I saw Colias palaeno and Plebejus optilete. Last confirmed sighting in Belgium was in 2006 so this species has most probably been lost.



Erebia meolans
On the high parts of the Black Forest this is one of the more common butterflies. Together with the High Vosges in neighboring France these are the north-westernmost populations of this species.

upperside male

underside female